There are two high-energy phosphate bond equivalents are used in the pathway leading from imp to gmp.
In the biosynthesis of gmp from imp, the conversion of imp + glutamine to gmp + glutamate requires the hydrolysis of atp to amp + ppi. The loss of pyrophosphate is equivalent to the loss of two high-energy phosphate bonds.
Imp is first converted to adenylosuccinate catalyzed by adenylosuccinate synthase. This is converted to amp by adenylosuccinate lyase. For conversion to gmp, imp is converted to xanthosine monophosphate by the action of imp dehydrogenase. Xanthosine monophosphate is converted to gmp by gmp synthase.
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