Check all that are true regarding the anatomy and physiology of the sense of equilibrium. Check All That Apply Otoliths are small calcium carbonate crystals that are embedded within a gelatinous layer. Otoliths are small calcium carbonate crystals that are embedded within a gelatinous layer. Otoliths and the gelatinous layer form the otolithic membrane. Otoliths and the gelatinous layer form the otolithic membrane. When the head is held erect, the otolithic membrane applies pressure directly on the hair cells. When the head is held erect, the otolithic membrane applies pressure directly on the hair cells. When the head is tilted, the hair cells remain erect, but there is a change in neurotransmitter release.